The Cretaceous pterosaur Tupandactylus navigans is known for having an anomalously oversized head crest. Until an almost complete fossil skeleton was found in Brazil, paleontologists had been able to study only skull specimens from T. navigans, though it was presumed that, like other pterosaurs, the species’s primary form of locomotion was powered flight. Examining the fuller skeleton in 2016, Victor Beccari and his team determined that T. navigans had long hind legs, short wings, and an unusually long neck—characteristics that, combined with the creature’s large-crested head, would have made sustained flight difficult and walking upright relatively comfortable. Based on these findings the team suggests that T. navigans likely ______blank