Birds of many species ingest foods containing carotenoids, pigmented molecules that are converted into feather coloration. Coloration tends to be especially saturated in male birds’ feathers, and because carotenoids also confer health benefits, the deeply saturated colors generally serve to communicate what is known as an honest signal of a bird’s overall fitness to potential mates. However, ornithologist Allison J. Shultz and others have found that males in several species of the tanager genus Ramphocelus use microstructures in their feathers to manipulate light, creating the appearance of deeper saturation without the birds necessarily having to maintain a carotenoid-rich diet. These findings suggest that ______blank